Philippine Mammal Project: Publications

PUBLICATIONS

The following is a list of publications about Philippine mammals since 1978 including publications associated with
the Philippine Mammal Project and other relevant research.
Last updated 1 September 2022
* = non-technical, not peer reviewed

2023

Balete, D. S., L. R. Heaney, E. A. Rickart, R. S. Quidlat, D. M. Rowsey and L. E. Olson. 2023. A re-assessment of diversity among Philippine gymnures (Mammalia: Erinaceidae: Podogymnura) with a new species from eastern Mindanao. Zootaxa, 5228 (3): 244–266.
Smith, S. M., D. M. Rowsey, J. A. Nations, K. D. Angielczyk, and L. R. Heaney. 2023. The roles of phylogeny, body size, and substrate use in vertebral trabecular bone variation in a radiation of small mammals. Biological Journal of the Linnean Society, 140 (1): 1-25. DOI: 10.1093/biolinnean/blad033.
Nachman, M. W., et al. (117 co-authors including L. R. Heaney). 2023. Specimen collection is essential for modern science. PLoS Biol 21(11): e3002318. doi.org/10.1371/journal.pbio.3002318
Heaney, L. R. 2024. Celebrating a life of undaunted curiosity. Review of Costa, J. T. 2023. Radical by Nature: the Revolutionary Life of Alfred Russel Wallace. Princeton University Press, Princeton. 515 pp. Evolution. DOI: 10.1093/evolut/qpad235

2022

Baletemys kampalili. Rowsey, Duya, Ibanez, Jansa, Rickart, & Heaney. Journal of Mammalogy, 103:xxx, DOI .org/10.1093/jmammal/gyac057.
Rowsey, D. M, M. R. M. Duya, J. C. Ibanez, S. A. Jansa, E. A. Rickart, and L. R.Heaney. 2022. A new genus and species of shrew-like mouse (Rodentia: Muridae) from a new center of endemism in eastern Mindanao, Philippines. Journal of Mammalogy, 103:1259–1277. doi.org/10.1093/jmammal/gyac057.
Ali, J. R., and L. R. Heaney. 2022. Alfred Russel Wallace and his enduring impact on biogeographical studies of the Indo-Australian Archipelago. Journal of Biogeography, 50:32-40. DOI:10.1111/jbi14470.
Heaney, L. R. 2022. Honoring Danny Balete, Philippine biodiversity scientist and conservationist. In the Field, 93: 21.

2021

Reginaldo, A. A., P. S. Ong, and L. R. Heaney. 2021. A report on the small mammals in mixed-use urbanized habitats in the southern Central Cordillera, northern Luzon. Philippine Science Letters, 14:282-290.
Petrosky, A. L., D. M. Rowsey, and L. R. Heaney. 2021. Molecular assessment of dietary niche partitioning in an endemic island radiation of tropical mammals. Molecular Ecology, https://doi.org/10.1111/mec.16158.
Heaney, L. R., C. C. Kyriazis, D. S. Balete, E. A. Rickart, J. M. Bates. 2021. A new species of Bullimus (Muridae, Rodentia) from southern Luzon Island, Philippines. Proceedings of the Biological Society of Washington, 134:131-148.
Ochoa, J., A. S. B. Mijares, P. J. Piper, M. C. Reyes, and L. R. Heaney. 2021. Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy, 102:909–930. DOI:10.1093/jmammal/gyab023.
Ali, J. R., and L. R. Heaney. 2021.Wallace’s Line, Wallacea, and associated divides and areas: a history of a tortuous tangle of ideas and labels. Biological Reviews, 96:922-942. DOI: 10.1002/BRV.12683.
Rickart, E. A., L. R. Heaney, and D. S. Balete. 2021. Mammals of Mt. Pintubo, Luzon Island, Philippines: extreme resilience following catastrophic disturbance. Philippine Journal of Science, 150:121-133.
Nations, J. A., A. Mursyid, R. D. Busta, S. P. Adrian, H. Handika, Apandi, A. S. Achmadi, and J. A. Esselstyn. 2021. The first report of albinism in a Sundaland endemic rodent. Mammalia, 85:168-172.
Nations, J. A., G. G. Mount, S. M. Morere, A. S. Achmadi, K. C. Rowe, and J. A. Esselstyn. 2021. Locomotory mode transitions alter phenotypic evolution and lineage diversification in an ecologically rich clade of mammals. Evolution, 75:376-393.
Reginaldo, A. A., K. C. Q. Soriano, B. B. Iglesia, and C. G. Vertudes. 2021. Occurrence and habitat use of non-native small mammal in upland urban-forest environment in Northern Philippines. Journal of Environmental Science and Management, 24-2: 62–74.
Reginaldo, A. A., and P. S. Ong. 2021. Small mammal–microhabitat associations in a disturbed landscape in northern Luzon, Philippines. Philippine Science Letters, 14(1): 51–64.
Sedlock, J. L., Gomes, D. G. E., Rubin, J. J., Woody, S., Hadi, B. A. R., and J. R. Barber. 2021. A phantom ultrasonic insect chorus repels low-flying bats, but most are undeterred. Functional Ecology, (April), 1–10.

2020

Balete, D. S., E. A. Rickart, and L. R. Heaney. 2020. A comparison of native and non-native small mammals in adjacent karst and volcanic regions in NE Luzon, Philippines. Philippine Journal of Systematic Biology, 14 (2):1-14.
Duya, M. R. M., L. R. Heaney, E. S. Fernando, and P. S. Ong. 2020. Fruit bat assemblages in different lowland forest types inn the Northern Sierra Madre Mountains, Philippines. Acta Chiropterologica, 22: 95-112.
Sedlock, J. S., L. R. Heaney, and M. Ruedi. 2020. Philippine bats of the genus Kerivoula (Chiroptera: Vespertilionidae): Overview and assessment of variation in K. pellucida and K. whiteheadi. Zootaxa, 4755 (3): 454–490.
Roycroft, E. J., J. A. Nations, and K. C. Rowe. 2020. Environment predicts repeated body size shifts in a recent radiation of Australian mammals. Evolution, 74:671-680.
Reginaldo A. A., and P. S. Ong. 2020. Structure of small non-flying mammal communities in disturbed habitats in the Central Cordillera, Luzon Island, Philippines. Philippine Science Letters, 13:81–94.
Nina R. Ingle, 2020. Indigenous People’s Education During COVID-19: An Environmentalist’s Perspective. Arete, Ateneo de Manila (web publication).*

2019

Nations, J. A., L. R. Heaney, T. C. Demos, A. S. Achmadi, K. C. Rowe, and J. A. Esselstyn. 2019. A simple skeletal measurement predicts climbing behavior in murine rodents. Biological Journal of the Linnean Society,128, 323–336.
Rickart, E. A., D. S. Balete, R. M. Timm, P. A. Alviola, J. A. Esselstyn, and L. R. Heaney. 2019. Two new species of shrew-rats (Rhynchomys: Muridae: Rodentia) from Luzon Island, Philippines. Journal of Mammalogy, 100: 1112-1129.
Rowsey, D.M., L. R. Heaney, and S. A. Jansa. 2019. Tempo and mode of mandibular shape and size evolution reveal contrasting support for incumbency effects in two clades of island-endemic rodents (Muridae: Murinae). Evolution, 73: 1411–1427.
Breed, W. G., H. Hassan, M. Gonzalez, H. J. McLennan, C. M. Leigh, and L. R. Heaney. 2019. Interspecific diversity of testes mass and sperm morphology in the Philippine chrotomyine rodents : Implications for differences in breeding systems across the species. Reproduction, Fertility, and Development, 31: 705-711.
Sedlock, J. L., Stuart, A. M., Horgan, F. G., Hadi, B., Jacobson, A. C., Alviola, P. A., and J. D. V. Alvarez. 2019. Local-scale bat guild activity differs with rice growth stage at ground level in the Philippines. Diversity, 11(9).
Dimaculangan, F. D., Como Jacobson, A., Alviola, P., Alvarez, J., and J. L. Sedlock. 2019. Seasonal emergence counts from a multispecies horseshoe bat (Chiroptera: Rhinolophidae) roost in the Philippines. Journal of Bat Research and Conservation, 12(1), 46–49.

2018

Gonzalez, J. C. T., C. A. A. Layusa-Oliveros, L. E. Afuang, M. R. M. Duya, L. R. Heaney, D. S. Balete, and 17 additional authors. 2018. Review and update of the 2004 national list of threatened terrestrial fauna of the Philippines. Sylvatrop, 28: 73-144.
Martinez, Q., R. Lebrun, A. S. Achmadi, J. A. Esselstyn, A. R. Evans, L. R. Heaney, R. Portela-Miguez, K. C. Rowe, and P.-H. Fabre. 2018. Convergent evolution of an extreme dietary specialization, the olfactory system of worm-eating rodents. Scientific Reports, 8:17806, DOI 10.1038/s41598-018-35827-0.
Heaney, L. R., and C.C. Kyriazis. 2018. Four mice species, with a common origin, have evolved on a small island. Natural History, 126: 12-15.*
Rowsey, D.M., L. R. Heaney, and S. A. Jansa. 2018. Diversification rates of the “Old Endemic” murine rodents of Luzon Island, Philippines are inconsistent with incumbency and ecological opportunity. Evolution, 72:1420-1435.
Heaney, L. R., Christopher C. Kyriazis, Danilo S. Balete, Scott J. Steppan, and Eric A. Rickart. 2018. How small an island? Speciation by endemic mammals (Apomys, Muridae) on an oceanic Philippine island. Journal of Biogeography, 45: 1675-1687.
Hutterer, R., D. S. Balete, T. C. Giarla, L. R. Heaney, and J. A. Esselstyn. 2018. A new genus and species of shrew (Mammalia: Soricidae) from Palawan, Philippines. Journal of Mammalogy, 99: 518–536.
Borgess, P., P. Cardoso, and 50 additional authors including L. R. Heaney. 2018. Global Island Monitoring Scheme (GIMS): a proposal for the long-term coordinated survey and monitoring of native island forest biota. Biodiversity and Conservation, 27: 2567-2586.
Kyriazis, C. C., B. Alam, M. Wjodyla, P. Hosner, S. Hackett, H.L. Mays Jr., L. R. Heaney, and S. Reddy. 2018. Colonization and diversification of the White-browed Shortwing (Aves: Muscicapidae: Brachypteryx montana) in the Philippines. Molecular Phylogeny and Evolution, 121: 121-131.

2017

Kyriazis, C., J. B. Bates, and L. R. Heaney. 2017. Dynamics of genetic and morphological diversification in an incipient intra-island radiation of Philippine rodents (Muridae: Bullimus). Journal of Biogeography, 44: 2585–2594.

2016

Heaney, L.R., D.S. Balete, S.A. Jansa, E.A. Rickart and S. Steppan. 2016. The role of topography in speciation by mammals within a complex plate-margin island. Pp. 409 in: R. Gabriel, R.B. Elias, I.R. Amorim & P.A.V. Borges (Eds). 2nd International Conference on Island Evolution, Ecology and Conservation: Island Biology 2016. Arquipelago. Life and Marine Sciences. Supplement 9. (Abstract only).*
Heaney, L. R., D. S. Balete, and E. A Rickart. 2016. Discovering diversity: Mammals of Luzon Island. In the Field, 87:10.*
Heaney, L. R., D. S. Balete, M. R. M. Duya, M. V. Duya, S. A. Jansa, S. J. Steppan, and E. A. Rickart. 2016. Doubling diversity: a cautionary tale of previously unsuspected mammalian diversity on a tropical oceanic island. Frontiers of Biogeography, 8.2, e29667, pp 1-19.
Dawson, M. N., A. C. Algar, L. R. Heaney, and Y. E. Stuart. 2016. The evolutionary biogeography of islands, lakes, and mountaintops. Pp. 203-210. In Encyclopedia of Evolutionary Biology (eds. R. M. Kliman and R. Gillespie). University of California Press, Berkeley.
Borregaard, M. K., I. R. Amorim, P. A. V. Borges, J. S. Cabral, J-M. Fernandez-Palacios, R. Field, L. R. Heaney, H. Kreft, T. J. Matthews, J. M. Olesen, J. Price, F. Rigal, M. J. Steinbauer, K. A. Triantis, L. Valente, P. Weigelt, and R. J. Whittaker. 2017. Oceanic island biogeography through the lens of the General Dynamic Model: assessment and prospect. Biological Reviews, 92: 830–853. DOI:10.1111/brv.12256.
Rickart, E. A., D. S. Balete, P. A. Alviola, M. J. Veluz, and L. R. Heaney. 2016. The mammals of Mt. Amuyao: a richly endemic fauna in the Central Cordillera of northern Luzon Island, Philippines. Mammalia, 80:579-592.
Heaney, L. R., D. S. Balete, and E. A. Rickart. 2016. The Mammals of Luzon Island: Biogeography and Natural History of a Philippine Fauna. Johns Hopkins University Press, Baltimore, xi + 287 pp.
Bailey, S. E., X. Mao, M. Bozek, G. Tsagkogeorga, G. Csorba, L. R. Heaney, J. Sedlock, W. Stanley, J-M. Rouillard, and S. J. Rossiter. 2016. Successful targeted sequence capture of degraded museum samples - a phylogenomic study of horseshoe bats. Biological Journal of the Linnean Society, 117: 58–70.

2015

Heaney, L. R. and D. S. Balete. 2015. Documenting and conserving biodiversity in the Philippines. In The Field, 86: 9.*
Balete, D. S., E. A. Rickart, L. R. Heaney, and S. A. Jansa. 2015. A new species of Batomys (Muridae, Rodentia) from Luzon Island, Philippines. Proceedings of the Biological Society of Washington, 128:22-39.
Justiniano, R., J. J. Schenk, D. S. Balete, E. A. Rickart, J. A. Esselstyn, L. R. Heaney, and S. J. Steppan. 2015. Testing diversification models of endemic Philippine forest mice (Apomys) with nuclear phylogenies across elevational gradients reveal repeated colonization of isolated mountain ranges. Journal of Biogeography, 42:51-64.

2014

Heaney, L. R., D. S. Balete, E. A. Rickart, and S. A. Jansa. 2014. Three new species of Musseromys (Muridae, Rodentia), the endemic Philippine tree mouse from Luzon Island. American Museum Novitates, 3802:1-28.
Heaney, L. R., D. S. Balete, M. J. Veluz, S. J. Steppan, J. A. Esselstyn, A. W. Pfieffer, and E. A. Rickart. 2014. Two new species of Philippine forest mice (Apomys, Muridae, Rodentia) from Lubang and Luzon Islands, with a redescription of Apomys sacobianus Johnson, 1962. Proceedings of the Biological Society of Washington, 126: 395-413.
Reginaldo, A.A., and AP.O. de Guia. 2014. Species richness and patterns of occurrence of small non-flying mammals of Mt. Sto. Tomas, Luzon Island, Philippines. Philippine Science Letters, 7(1):34–44.
Sedlock, J. L., Krüger, F., and E. L. Clare. 2014. Island bat diets: Does it matter more who you are or where you live? Molecular Ecology, 23(15).
Sedlock, J. L., Jose, R. P., Vogt, J. M., Paguntalan, L. M. J., and A. B. Cariño. 2014. A survey of bats in a karst landscape in the central Philippines. Acta Chiropterologica, 16(1), 197–211.

2013

Balete, D. S., L. R. Heaney, P. A. Alviola, and Eric A. Rickart. 2013. Diversity and distribution of small mammals in the Bicol Volcanic Belt of southern Luzon Island, Philippines. National Museum of the Philippines Journal of Natural History, 1:61-86.
Heaney, L. R., D. S. Balete, R. G. B. Rosell Ambal, M. J. Veluz, and E. A. Rickart. 2013. The non-volant small mammals of Mt. Banahaw - San Cristobal National Park, Luzon, Philippines: Distribution and ecology of a highly endemic fauna. National Museum of the Philippines Journal of Natural History, 1:45-60.
Rickart, E. A., L. R. Heaney, D. S. Balete, P. A. Alviola, M. R. M. Duya, M. V. Duya, G. Rosell-Ambal, and J. L. Sedlock. 2013. The mammals of Mt. Natib, Bataan Province, Luzon, Philippines. National Museum of the Philippines Journal of Natural History, 1:31-44.
Balete, D. S., L. R. Heaney, and E. A. Rickart. 2013. The mammals of Mt. Irid, Southern Sierra Madre, Luzon Island. National Museum of the Philippines Journal of Natural History, 1:15-29.
Heaney, L. R., D. S. Balete, P. A. Alviola, M. R. M. Duya, and E. A. Rickart. 2013. The mammals of Mt. Anacuao, NE Luzon Island, Philippines: A test of predictions of Luzon small mammal diversity patterns. National Museum of the Philippines Journal of Natural History, 1:1-13.
Heaney, L. R. 2013. Preface to studies of mammalian biodiversity on Luzon Island, Philippines. National Museum of the Philippines Journal of Natural History, 1:iii-v.
Heaney, L. R., D. S. Balete, and E. A. Rickart. 2013. Models of oceanic island biogeography: changing perspectives on biodiversity dynamics in archipelagoes. Frontiers of Biogeography, 5:249-257.
Lorica, M. R. P., and L. R. Heaney. 2013. Survival of a native mammalian carnivore, the leopard cat Prionalurus bengalensis Kerr, 1792 (Carnivora: Felidae), in an agricultural landscape on an oceanic Philippine island. Journal of Threatened Taxa, 5:4451-4560.
Heaney, L. R. 2013. Introduction and Commentary. Pp. xi-lxxi. In A. R. Wallace, Island Life (reprint of 1880 first edition). University of Chicago Press, Chicago.
Fernando, E.S. and Balete, D.S. 2013. Medinilla dallciana (Melastomatacea: Dissochaeteae), a new species from Luzon Islands, Philippines. Philippine Journal of Science, 142: 89-93.

2012

Heaney, L. R., D. S. Balete, P. A. Alviola, E. A. Rickart, and M. Ruedi. 2012. Nyctalus plancyi and Falsistrellus petersi (Chiroptera: Vespertillionidae) from northern Luzon, Philippines: ecology, phylogeny, and biogeographic implications. Acta Chiropterologica, 14: 265-278.
Heaney. L. R. 2012. Shrinking giants. In the Field, 83:12.*
Balete, D. S., E. A. Rickart, L. R. Heaney, P. A. Alviola, M. R. M. Duya, M. V. Duya, T. Sosa, and S. A. Jansa. 2012. Archboldomys (Muridae: Murinae) reconsidered: A new genus and three new species of shrew mouse from Luzon Island, Philippines. American Museum Novitates, 3754: 1-60.
Esselstyn, J. A., J. L. Sedlock, F. A. Anwarali Khan, B. J. Evans, and L. R. Heaney. 2012. Single-locus species delimitation: A test of the mixed Yule-coalescent model, with an empirical application to Philippine round-eared bats. Proceedings of the Royal Society B, 279: 3678-3686.
Heaney, L. R., and D. S. Balete. 2012. Discovering diversity: newly discovered mammals highlight areas of unique biodiversity in Luzon. Wildernews, 1:4-9.*

2011

Heaney, L. R., P. J. Piper, and A. S. Mijares. 2011. The first fossil record of endemic murid rodents from the Philippines: A Late Pleistocene cave fauna from northern Luzon. Proceedings of the Biological Society of Washington, 124:234-247.
Duya, M. R. M., M. V. Duya, P. A. Alviola, D. S. Balete, and L. R. Heaney. 2011. Diversity of small mammals in montane and mossy forests on Mount Cetaceo, Cagayan Province, Luzon. Fieldiana Life and Earth Sciences, 2:88-95.
Balete, D. S., P. A. Alviola, M. R. M. Duya, Melizar V. Duya, L. R. Heaney, and E. A. Rickart. 2011. The mammals of the Mingan Mountains, Luzon: evidence for a new center of mammalian endemism. Fieldiana Life and Earth Sciences, 2: 75-87.
Alviola, P. A., M. R. M. Duya, M. V. Duya, L. R. Heaney, and E. A. Rickart. 2011. Mammalian diversity patterns on Mt. Palali, Caraballo Mountains, Luzon. Fieldiana Life and Earth Sciences, 2:61-74.
Heaney, L. R., E. A. Rickart, D. S. Balete, M. V. Duya, M. R. M. Duya, and S. Steppan. 2011. Seven new species and a new subgenus of forest mice (Rodentia: Muridae: Apomys) from Luzon Island. Fieldiana Life and Earth Sciences, 2:1-60.
Heaney, L. R. (ed.). 2011. Discovering Diversity: Studies of the Mammals of Luzon Island, Philippines. Fieldiana Life and Earth Sciences, 2:vii + 102 pp.
Heaney, L. R. 2011. Island biogeography: Paradigm lost? Frontiers of Biogeography, 2:94-97.
Rickart, E. A., D. S. Balete, R. J. Rowe, and L. R. Heaney. 2011. Mammals of the northern Philippines: tolerance for habitat disturbance and resistance to invasive species in an endemic fauna. Diversity and Distributions, 17:530-541.
Rickart, E. A., L. R. Heaney, D. S. Balete, and B. R. Tabaranza, Jr. 2011. Small mammal diversity along an elevational gradient in northern Luzon, Philippines. Mammalian Biology, 76: 12-21.
Barcelona, J.F., E.S. Fernando, D.L. Nickrent, D.S. Balete & P.B. Pelser. 2011. An amended description of Rafflesia leonardi and a revised key to Philippine Rafflesia (Rafflesiaceae). Phytotaxa, 24: 11-18.
Sedlock, J.L., N.R. Ingle & D.S. Balete. 2011. Enhanced sampling of bat assemblages: a field test on Mount Banahaw, Luzon. Fieldiana: Life and Earth Sciences, 2: 96–102.

2010

Hoffmann, M., C. Hilton-Taylor, A. Angulo, et al. (206 authors, including L. R. Heaney). 2010. The impact of conservation on the status of the world’s vertebrates. Science, 330:1503-1509.
Esselstyn, J. A. & S. M. Goodman. 2010. New species of shrew (Soricidae: Crocidura) from Sibuyan Island, Philippines Journal of Mammalogy 91:1467–1472.
Heaney, L. R. 2010. (Review of) The Theory of Island Biogeography Revisited. Quarterly Review of Biology, 85:350-351.
Heaney, L. R., M. L. Dolar, D. S. Balete, J. A. Esselstyn, E. A. Rickart, and J. L. Sedlock. 2010. Synopsis of Philippine Mammals. Field Museum website, http://www.fieldmuseum.org/philippine_mammals/.
Brown, R.M., C.W. Linkem, A.C. Diesmos, D.S. Balete, M.V. Duya & J. Ferner. 2010. Species boundaries in Philippine montane forest Sphenomorphus: three new species from the mountains of Luzon and clarification of the status of the poorly-known S. beyeri, and S. knollmanae and S. laterimaculatus. Scientific Papers, Natural History Museum, University of Kansas, 42:1–27.
Balete, D.S., P.B. Pelser, D.L. Nickrent & J.F. Barcelona. 2010. Rafflesia verrucosa (Rafflesiaceae), a new species of small-flowered Rafflesia from Mindanao, Philippines. Phytotaxa, 10: 49–57.

2009

Heaney, L. R., and T. E. Roberts. 2009. New perspectives on the long-term biogeographic dynamics and conservation of Philippine fruit bats. Pp. 17-58. In T. H. Fleming and P. Racey (eds.), Ecology, Evolution, and Conservation of Island Bats. University of Chicago Press.
Heaney, L. R., D. S. Balete, E. A. Rickart, M. J. Veluz, and S. Jansa. 2009. A new genus and species of small “tree mouse” (Rodentia, Muridae) related to the Philippine giant cloud-rats. In R. S. Voss and M. D. Carleton (eds.), Systematic Mammalogy, Contributions in Honor of Guy G. Musser. Bulletin of the American Museum of Natural History, 331: 205-229.
Balete, D. S., L. R. Heaney, M. J. Veluz, and E. A. Rickart. 2009. Diversity patterns of small mammals in the Zambales Mts., Luzon, Philippines. Mammalian Biology, 74:456-466.
Heaney, L. R., and M. V. Lomolino. 2009. From the foundations to the frontiers of biogeography. Frontiers of Biogeography, 1:3-4.
Heaney, L. R., and S. M. Goodman. 2009. Mammal radiations. Pp. 588-591. In R. Gillespie and D. Clague (eds.), Encyclopedia of Islands. University of California Press.
Danielsen, F., N.D. Burgess, A. Balmford, P.F. Donald, M. Funder, J.P.G. Jones, P. Alviola, D.S. Balete, T. Blomley, J. Brashares, B. Child, M. Enghoff, J. Fjeldså, S. Holt, H. Hübertz, A.E. Jensen, P.M. Jensen, J. Massao, M.M. Mendoza, Y. Ngaga, M.K. Poulsen, R. Rueda, M. Sam, T. Skielboe, G. Stuart-Hill, E. Topp-Jørgensen and D. Yonten. 2009. Local participation in natural resource monitoring: a characterization of approaches. Conservation Biology, 23: 31–42.
Barcelona, J.F., P.B. Pelser, D.S. Balete, and L.L. Co. 2009. Taxonomy, ecology, and conservation status of Philippine Rafflesia (Rafflesiaceae). Blumea, 54: 77–93.
Barcelona, J.F., L.L. Co, D.S. Balete, and N.A. Bartolome. 2009. Rafflesia aurantia (Rafflesiaceae): a new species from northern Luzon, Philippines. Gardens’ Bulletin Singapore, 61: 17–27.
Gonzales, R. S., N.R. Ingle, D.A. Lagunzad, and T. Nakashizuka. 2009. Seed dispersal by birds and bats in lowland Philippine forest successional area. Biotropica, 41: 452–458.

2008

Duya, M. R. M., M. V. Duya, P. A. Alviola, D. S. Balete, and L. R. Heaney. 2008. Report on a Survey of the Mammals of the Sierra Madre Range, Luzon Island, Philippines. Banwa, 4 (2007):41-68.
Ong, P. S, and E. A. Rickart. 2008. Ecology of native and pest rodents in the Philippines. Pp 1-15 in
Philippine Rats: Ecology and Management (G. R. Singleton, R. C. Joshi, and L. S. Sebastian, eds), Philippine Rice Research Institute.
Balete, D. S., L. R. Heaney, E. A. Rickart, R. S. Quidlat, and J. C. Ibanez. 2008. A new species of Batomys (Muridae: Murinae) from eastern Mindanao Island, Philippines. Proceedings of the Biological Society of Washington, 121: 411-428.
Jan Schipper and 131 other authors, including Heaney, L. R. 2008. Status of the World’s Land and Marine Mammals: Diversity, Threat, and Knowledge. Science, 322: 225-230.
Esselstyn, J. A., H. J. D. Garcia, M. G. Saulog, and L. R. Heaney. 2008. A new species of Desmalopex (Pteropodidae) from the Philippines, with a phylogenetic analysis of the pteropodini. Journal of Mammalogy, 89: 815-825.
Sedlock, J. L., Weyandi, S. E., Cororan, L., Damerow, M., Hwa, S. H., and B. Pauli. 2008. Bat diversity in tropical forest and agro-pastoral habitats within a protected area in the Philippines. Acta Chiropterologica, 10(2), 349–358.

2007

Rickart, E. A., D. S. Balete, and L. R. Heaney. 2007. Habitat disturbance and the ecology of small mammals in the Philippines. Journal of Environmental Science and Management, 10: 34-41.
Heaney, L. R. 2007. Islands Rising (Review of Island Biogeography: Ecology, Evolution, and Conservation, 2nd edition, but R. J. Whittaker and J. M. Fernandez-Palacios). Journal of Biogeography, 34: 1832.
Balete, D. S., E. A. Rickart, R. G. B. Rosell-Ambal, S. Jansa, and L. R. Heaney. 2007. Descriptions of two new species of Rhynchomys Thomas (Rodentia: Muridae: Murinae), from Luzon Island, Philippines. Journal of Mammalogy, 88: 287-301.
Heaney, L. R. 2007. Is a new paradigm emerging for oceanic island biogeography? Journal of Biogeography, 34: 753-757.
Danielsen, F., M. M. Mendoza, A. Tagtag, P. A. Alviola, D. S. Balete, A. E. Jensen, M. Enghoff & M.K. Poulsen. 2007. Increasing conservation management action by involving local people in natural resource management. Ambio, 36: 566–570.
Helgen, K. M., D. Kock, R. K. S. C. Gomez, N. R. Ingle, and M.H. Sinaga. 2007. Taxonomy and natural history of the Southeast Asian fruit-bat genus Dyacopterus. Journal of Mammalogy 88(2):302-318.

2006

Heaney, L. R., D. S. Balete, J. Sarmiento, and P. A. Alviola. 2006. Losing diversity and courting disaster: The mammals of Mt. Data National Park. Haring Ibon 25: 15-23.*
Heaney, L. R., B. R. Tabaranza Jr., D. S. Balete, E. A. Rickart, and N. R. Ingle. 2006. The mammals of Mt. Kitanglad Nature Park, Mindanao, Philippines. Fieldiana: Zoology, new series, 112: 1-63.
Balete, D. S., E. A. Rickart, and L. R. Heaney. 2006. A new species of the shrew-mouse, Archboldomys (Rodentia: Muridae: Murinae) from the Philippines. Systematics and Biodiversity, 4:489-501.
Croft, D. A., L. R. Heaney, J. J. Flynn, and A. P. Bautista. 2006. Fossil remains of a new, diminutive Bubalus (Artiodactyla: Bovidae: Bovini) from Cebu Island, Philippines. Journal of Mammalogy, 87:1037-1051.
Balete, D. S., B. R. Tabaranza, Jr., and L. R. Heaney. 2006. An annotated checklist of the land birds of Camiguin Island, Philippines. Fieldiana Zoology new series, 106:58-72.
Heaney, L. R., B. R. Tabaranza, Jr., D. S. Balete, and N. Rigertas. 2006. Synopsis and biogeography of the mammals of Camiguin Island, Philippines. Fieldiana Zoology new series, 106:28-48.
Heaney, L. R. and B. R. Tabaranza, Jr. 2006. A new species of forest mouse, genus Apomys (Mammalia: Rodentia: Muridae) from Camiguin Island, Philippines. Fieldiana Zoology, new series, 106:14-27.
Heaney, L. R. and B. R. Tabaranza, Jr. 2006. Mammal and Land Bird Studies on Camiguin Island, Philippines: Background and Conservation Priorities. Fieldiana Zoology, new series, 106:1-13.
Heaney, L. R. (editor). 2006. The Mammals and Birds of Camiguin Island, Philippines, an Distinctive Center of Biodiversity. Fieldiana Zoology, new series, 106:1-72.
Catibog-Sinha, C. C., and L. R. Heaney. 2006. Philippine Biodiversity, Principles and Practice. Haribon Foundation, Philippines. 495 pp.
Jansa, S., K. Barker, and L. R. Heaney. 2006. The pattern and timing of diversification of Philippine endemic rodents: Evidence from mitochondrial and nuclear gene sequences. Systematic Biology, 55:73-88.
Balete, D.S., R.S. Quidlat & J.C. Ibanez. 2006. Non-volant mammals of Mt. Hamiguitan, eastern Mindanao, Philippines. Banwa, 3: 65 – 80.

2005

Heaney, L. R. 2005. A second chance to avert extinction: good news for Philippine biodiversity. Silliman Journal, 45 (2004):111-112.*
Balete, D. S., L. R. Heaney, J. Sarmiento, and M. J. Veluz. 2005. Zambales surprises: The mammals of Mt. Tapulao. Haring Ibon, 21:20-27.*
Rickart, E. A., L. R. Heaney, S. M. Goodman, and S. Jansa. 2005. Review of the Philippine genera Chrotomys and Celaenomys (Murinae) and description of a new species. Journal of Mammalogy, 86:415-428.
Heaney, L. R., D. S. Balete, G. V. Gee, M. V. Lepiten-Tabao, E. A. Rickart, and B. R. Tabaranza. 2005. Preliminary report on the mammals of Balbalasang, Kalinga Province, Luzon. Sylvatrop, 13 (2003):51-62.
Heaney, L. R. 2005. Remarkable rats. USA Today, 133:44-47.*
Heaney, L. R., Walsh, J. S. Jr., and A. T. Peterson. 2005. The roles of geological history and colonization abilities in genetic differentiation between mammalian populations in the Philippine archipelago. Journal of Biogeography, 32:229-247.
Danielsen, F., A.E. Jensen, P.A. Alviola, D.S. Balete, M.M. Mendoza, A. Tagtag, C. Custudio & M. Enghoff. 2005. Does monitoring matter? A quantitative assessment of management decisions from locally-based monitoring of protected areas. Biodiversity and Conservation, 14: 2633 – 2652.

2004

Esselstyn, J. A., P. Widmann & L. R. Heaney. 2004. The Mammals of Palawan Island, Philippines. Proceedings of the Biological Society of Washington, 117:271-302.
Heaney, L. R. 2004. Conservation Biogeography in an Oceanic Archipelago. Pp. 345-360. In M. V. Lomolino and L. R. Heaney (eds.), Frontiers of Biogeography, New Directions in the Geography of Nature. Sinauer Associates, Sunderland.
Lomolino, M. V., and L. R. Heaney (eds.). 2004. The Frontiers of Biogeography: New Directions in the Geography of Nature. Sinauer Associates, Sunderland.
Heaney, L. R., D. S. Balete, E. A. Rickart, M. J. Veluz, and J. Sarmiento. 2004. Welcome surprises from Mt. Banahaw. Haring Ibon, 16:10-15.*
Heaney, L. R. and G. Vermeij. 2004. Diversification. Pp. 779-788. In M. V. Lomolino, J. H. Brown, and D. Sax (eds.). Foundations of Biogeography. University of Chicago Press, 1291 pp.
Heaney, L. R. 2004. Remarkable rats and the origins of island biological diversity. In The Field, 75:16-18.*

2003

Steppan, S., C. Zawadski, and L. R. Heaney. 2003. Molecular phylogeny of the endemic Philippine rodent Apomys (Muridae) and the dynamics of diversification in an oceanic archipelago. Biological Journal of the Linnean Society, 80:699-715.
Rickart, E. A., L. R. Heaney, and B. R. Tabaranza, Jr. 2003. A new species of Limnomys (Rodentia: Muridae: Murinae) from Mindanao Island, Philippines. Journal of Mammalogy, 84:1443-1455.
Heaney, L. R., D. S. Balete, Genevieve V. A. Gee, and E. A. Rickart. 2003. Remarkable rats of the Central Cordillera. Haring Ibon, 13:18-29.*
Rickart, E. A. 2003. Chromosomes of Philippine mammals (Insectivora, Dermoptera, Primates, Rodentia, Carnivora). Proceedings of the Biological Society of Washington, 116:699-709.
Danielsen, F., M. M. Mendoza, A. Tagtag, P. A. Alviola, D. S. Balete, M. Enghoff, M. K. Poulsen and A. E. Jensen. 2003. On participatory biodiversity monitoring and its applicability – a reply to Yoccoz et al.& Rodriguez. Oryx, 37: 412.
Danielsen, F., M.M. Mendoza, P.A. Alviola, D.S. Balete, M. Enghoff, M.K. Poulsen & A.E. Jensen. 2003. Biodiversity monitoring in developing countries: what are we trying to achieve? Oryx, 37: 407 – 409.
Heideman, Paul D., and Utzurrum, Ruth C. 2003. Seasonality and synchrony of reproduction in three species of nectarivorous Philippines bats. BMC Ecology, 2003 Nov 21; 3(1): 11
Ingle, N. R. 2003. Seed dispersal by wind, birds, and bats between Philippine montane rainforest and successional vegetation. Oecologia, 134:251-261.

2002

Heaney, L. R. 2002. Biological Diversity in the Philippines: an Introduction to Megadiversity in a Nation of Islands. Pp 2-8. In P. S. Ong, L. E. Afuang, R. G. Rosell-Amball (eds.), Philippine Biodiversity Conservation Priorities. Department of Environment and Natural Resources and Conservation International Philippines, Quezon City, Pp xvii + 113.
Rickart, E. A., and L. R. Heaney. 2002. Further studies on the chromosomes of Philippine rodents (Muridae: Murinae). Proceedings of the Biological Society of Washington, 115:473-487.
Heaney, L. R. 2002. A decade of research on Philippine mammals: Progress and challenges. Silliman Journal, 42 (2001):88-108.
Rickart, E. A., L. R. Heaney, and B. R. Tabaranza, Jr. 2002. Review of Bullimus (Muridae: Murinae) and description of a new species from Camiguin Island, Philippines. Journal of Mammalogy 83:421-436.
Heaney, L. R. and N. A. D. Mallari. 2002. A preliminary analysis of current gaps in the protection of threatened Philippine terrestrial mammals. Sylvatrop, 10 (2000):28-39.
Heaney, L. R., E. K. Walker, B. R. Tabaranza, Jr., and N. R. Ingle. 2002. Mammalian diversity in the Philippines: an assessment of the adequacy of current data. Sylvatrop, 10 (2000):6-27.
Heaney, L. R. 2002. Island life along Wallace’s Line: Biogeography and patterns of endemism in the Philippines and Indonesia. Pp. 28-30. In Wikramanayake, E., E. Dinerstein, C.J. Loucks, D.M. Olson, J. Morrison, J. Lamoreaux, M. McKnight, and P. Hedao. Terrestrial ecoregions of the Indo-Pacific: A conservation assessment. Washington, DC: Island Press, 643 pp.
Sedlock, J. L.. 2002. Autecology and the conservation of insectivorous bats on Mt. Makiling. Silliman Journal, 42(1), 163–201.

2001

Heaney, L. R. 2001. Small mammal diversity along elevational gradients in the Philippines: an assessment of patterns and hypotheses. Global Ecology and Biogeography, 10:15-39.
Heaney, L. R., M. V. Lomolino, and R. J. Whittaker (eds.) 2001. Diversity patterns of small mammals along elevational gradients. Special issue, Global Ecology and Biogeography, 10:1-109.
Sedlock, J. L.. 2001. Inventory of insectivorous bats on Mount Makiling, Philippines using echolocation call signatures and a new tunnel trap. Acta Chiropterologica, 3(2), 163–178.

2000

Heaney, L. R., A. Diesmos, B. R. Tabaranza Jr., N. A. Mallari, and R. Brown. 2000. Beacon of Hope - a first report from Kalinga Province, in the Northern Central Cordillera. Haring Ibon (Manila), 2:14-18.*
Heaney, L. R. 2000. Dynamic disequilibrium: a long-term, large-scale perspective on the equilibrium model of island biogeography. Global Ecology and Biogeography, 9:59-74.
Danielsen, F., D.S. Balete, M.K. Poulsen, M. Enghoff, C.M. Nozawa & A.E. Jensen. 2000. A simple system for monitoring biodiversity in protected areas of a developing country. Biodiversity Conservation, 9:1671–1705.

1990's

Heaney, L. R., N. R. Ingle, J. L. Sedlock, and B. R. Tabaranza, Jr. 1999. Non-flying mammals of Mindanao Island, Philippines. Laminated color field guide, 2pp. The Field Museum, Chicago.*
Ingle, N. R., J. L. Sedlock, and L. R. Heaney. 1999. Bats of Mindanao Island, Philippines. Laminated color field guide, 2pp. The Field Museum, Chicago.*
Heaney, L. R., P. S. Ong, R. A. Mittermeier, and C. G. Mittermeier. 1999. The Philippines. Pp. 308-317. In R. A. Mittermeier, N. Myers, P. Robles G., and C. G. Mittermeier (eds.), Hotspots, earth’s biologically richest and most endangered terrestrial ecosystems. CEMEX, Mexico City, 431 pp.*
Heaney, L. R. 1999. Historical biogeography in SE Asia: Integrating paradigms and refining the details. (Review of “Biogeography and Geological Evolution in Southeast Asia”, edited by R. Hall and J. D. Holloway). Journal of Biogeography, 26: 435-437.
Heaney, L. R., D. S. Balete, E. A. Rickart, R. C. B. Utzurrum, and P. C. Gonzales. 1999. Mammalian diversity on Mt. Isarog, a threatened center of endemism on southern Luzon Island, Philippines. Fieldiana Zoology, new series, 95:1-62.
Rickart, E. A., J. A. Mercier, and L. R. Heaney. 1999. Cytogeography of Philippine bats. Proceedings of the Biological Society of Washington, 112:453-469.
Heaney, L. R. 1999. Monitoring mammalian diversity in the Philippines: a discussion of issues. Sylvatrop (1997), 7:108 (abstract only).
Heaney, L. R. 1999. A Philippine Journey: the evolution of an environmental crisis. In the Field, 70:2-5.*
Musser, G. G., L. R. Heaney, and B. R. Tabaranza, Jr. 1998. Philippine rodents: redefinition of known species of Batomys (Muridae, Murinae) and description of a new species from Dinagat Island, Philippines. American Museum Novitates, 3237:1-51.
Rickart, E. A., L. R. Heaney, D. S. Balete, and B. R. Tabaranza, Jr. 1998. A review of the genera Crunomys and Archboldomys (Rodentia, Muridae, Murinae) with descriptions of two new species from the Philippines. Fieldiana Zoology new series, 89:1-24.
Heaney, L. R., D. S. Balete, M. L. Dolar, A. C. Alcala, A. T. L. Dans, P. C. Gonzales, N. R. Ingle, M. V. Lepiten, W. L. R. Oliver, P. S. Ong, E. A. Rickart, B. R. Tabaranza, Jr., and R. C. B. Utzurrum. 1998. A synopsis of the mammalian fauna of the Philippine Islands. Fieldiana Zoology, new series, 88:1-61.
Heaney, L. R. and J. C. Regalado, Jr. 1998. Vanishing Treasures of the Philippine Rain Forest. The Field Museum, Chicago. 88 pp.*
Heideman, Paul D., and Kendall S. Powell *. 1998. Age-specific reproductive strategies and delayed embryonic development in an Old World fruit bat, Ptenochirus jagori (Megachiroptera). Journal of Mammalogy 79:295-311. (Both authors contributed equally to this work; order of authorship was determinedby coin flip.)
Balete, D. S., and L. R. Heaney. 1997. Density, biomass, and movement estimates for murid rodents in mossy forest on Mount Isarog, southern Luzon, Philippines. Ecotropica, 3:91-100.
Heaney, L R. and R. A. Mittermeier 1997. The Philippines. In: R. A. Mittermeier, P.Robles Gil and C.G. Mittermeier (Eds.). Megadiversity. Earth's Biologically Wealthiest Nations. CEMEX, Monterrey, Mexico, pp. 236-255.*
Heaney, L. R., D. S. Balete, and A. T. L. Dans. 1997. Terrestrial Mammals. Pp. 116-144. In Wildlife Conservation Society of the Philippines, Philippine Red Data Book. Bookmark, Manila, Philippines. 240 pp.
Oliver, W. L. R., and L. R. Heaney. 1997. Biodiversity and conservation in the Philippines: an introduction to a global priority. Pp. xi-xx. In Wildlife Conservation Society of the Philippines, Philippine Red Data Book. Bookmark, Manila, Philippines. 240 pp.
Heaney, L. R. 1997. Current problems and research priorities for the conservation of mammalian diversity in the Philippines. Sylvatrop (1995), 5:87-88.
Heaney, L. R., and B. R. Tabaranza, Jr. 1997. A preliminary report on mammalian diversity and conservation status of Camiguin Island, Philippines. Sylvatrop, (1995), 5:57-64.
Heaney, L. R., C. R. Banks, L. E. Afuang, B. R. Tabaranza, Jr., and D. S. Balete (eds.). 1997. Fourth Annual Symposium - Workshop of the Wildlife Conservation Society of the Philippines on “Conservation of Threatened Species of Philippine Wildlife”. Sylvatrop, (1995), 5 (1 & 2):1-137.
Samson, D. A., E. A. Rickart, and P. C. Gonzales. 1997. Ant diversity and abundance along an elevational gradient in the Philippines. Biotropica, 29:349-363.
Oliver, W. L. R and L. R. Heaney. 1996. Biodiversity and conservation in the Philippines. International Zoo News, 43:329-337.
Custodio, C. C., M. V. Lepiten, and L. R. Heaney. 1996. Bubalus mindorensis. Mammalian Species, 520:1-5.
Balete, D. S., L. R. Heaney, and R. I. Crombie. 1995. First records of Hipposideros lekaguli Thonglongya and Hill 1974 from the Philippines. Asia Life Sciences, 4:89-94.
Heideman, P. D. 1995. Synchrony and seasonality of reproduction in tropical bats. pp 151 - 165 in Symposium of the Zoological Society of London No.67: Ecology, Evolution,and Behaviour of Bats. (P.A. Racey and S. Swift, Eds.) Oxford University Press.
Heaney, L. R. and M. Ruedi. 1994. A preliminary analysis of biogeography and phylogeny of shrews (Mammalia: Soricidae) from the Philippine Islands. pp 357-377. The biology of shrews; J. Merritt, G. Kirkland, and R. Rose, eds. Special Publications, Carnegie Museum.
Heaney, L. R. 1993. Biodiversity patterns and the conservation of mammals in the Philippines. Asia Life Sciences, 2:261-274.
Rickart, E. A., and G. G. Musser. 1993. Philippine rodents: chromosomal characteristics and their significance for phylogenetic inference among 13 species (Rodentia: Muridae: Murinae). American Museum Novitates, 3064:1-34.
Rickart, E. A. 1993. Diversity patterns of mammals along elevational and disturbance gradients in the Philippines: implications for conservation. Asia Life Sciences, 2:251-260.
Heaney, L. R., W. Gruezo, C. C. Custodio, and E. A. Rickart (eds.). 1993. Symposium on Conservation of Philippine Vertebrates. Asia Life Sciences, 2:101-304.
Peterson, A. T., and L. R. Heaney. 1993. Genetic differentiation in Philippine bats of the genera Cynopterus and Haplonycteris. Biological Journal of the Linnean Society, 49: 203-218.
Rickart, E. A., L. R. Heaney, P. D. Heideman, and R. C. B. Utzurrum. 1993. The distribution and ecology of mammals on Leyte, Biliran, and Maripipi islands, Philippines. Fieldiana Zoology, new series, 72:1-62.
Ingle, N. R. 1993. Vertical stratification of bats in a Phlippine rain forest. Asia Life Sciences, 2:215-222.
Heideman, P. D., J. A. Cummings, and L. R. Heaney. 1993. Observations on reproductive timing and early embryonic development in an Old World fruit bat, Otopteropus cartilagonodus (Megachiroptera). Journal of Mammalogy, 74:621-630.
Hoffmann, R. S., C. G. Anderson, R. W. Thorington, Jr., and L. R. Heaney. 1993. Family Sciuridae. Pp. 419-465, in Mammal Species of the World, a taxonomic and geographic reference, Second ed. (D. Wilson and D. M. Reeder, eds.). Smithsonian Institution Press, Washington, D.C. 1206 pp.
Oliver, W. L. R., C. R. Cox, P. C. Gonzales, and L. R. Heaney. 1993. Cloud rats in the Philippines - preliminary report on distribution and status. Oryx, 27:41-48.
Ingle, N. R. 1993. Vertical stratification of bats in a Philippine rain forest. Asia Life Sciences, 2:215‑222.
Goodman, S. M. and N. R. Ingle. 1993. Sibuyan Island in the Philippines — threatened and in need of conservation. Oryx, 27:174‑180.
Balete, D. S., H. C. Miranda, L. R. Heaney, and J. F. Rieger. 1992. Diversity and conservation of Philippine land vertebrates: an annotated bibliography. Silliman Journal, 36:129-149.
Heideman, P. D., and L. R. Heaney. 1992. (Authors of 20 species accounts in) Old-World Fruit Bats: An Action Plan for the Family Pteropodidae (S. P. Mickleburgh, P. A. Racey, and A. M. Hutson, eds.). IUCN Press.
Ingle, N. R. and L. R. Heaney. 1992. A key to the bats of the Philippine Islands. Fieldiana Zoology, new series, 69:1-44.
Musser, G. G., and L. R. Heaney. 1992. Philippine rodents: definitions of Tarsomys and Limnomys plus a preliminary assessment of phylogenetic patterns among native Philippine murines (Murinae, Muridae). Bulletin of the American Museum of Natural History, 211:1-138.
Heaney, L. R., and R. C. B. Utzurrum. 1991 (1992). A review of the conservation status of Philippine land mammals. Association of Systematic Biologists of the Philippines Communications, 3:1-13.
Heideman, P. D., and K. E. Erickson.1992. A new task for an old tool: Using crochet hooks to remove bats from mist nets. Bat Research News 33:7.
Ingle, N. R. 1992. The natural history of bats on Mt. Makiling, Luzon Island, Philippines. Silliman Journal, 36:1‑26.
Heaney, L. R. 1991. An analysis of patterns of distribution and species richness among Philippine fruit bats (Pteropodidae). Bulletin of the American Museum of Natural History, 206:145-167.
Heaney, L. R. 1991. A synopsis of climatic and vegetational change in Southeast Asia. Climatic Change, 19:53-61.
Rickart, E. A., L. R. Heaney, and R. B. Utzurrum. 1991. Distribution and ecology of small mammals along an elevational transect in southeastern Luzon, Philippines. Journal of Mammalogy, 72:458-469.
Heaney, L. R., P. C. Gonzales, R. C. B. Utzurrum, and E. A. Rickart. 1991. The mammals of Catanduanes Island: implications for the biogeography of small land-bridge islands in the Philippines. Proceedings of the Biological Society of Washington, 104:399-415.
Rickart, E. A., and L. R. Heaney. 1991. A new species of Chrotomys (Muridae) from Luzon Island, Philippines. Proceedings of the Biological Society of Washington, 104:387-398.
Utzurrum, R. C. B., and P. D. Heideman.1991. Differential injestion of viable vs. nonviable Ficus seeds by fruit bats. Biotropica 23:311-312.
Heaney, L. R., and E. A. Rickart. 1990. Correlations of clades and clines: geographic, elevational, and phylogenetic distribution patterns among Philippine mammals. pp. 321-332. In G. Peters and R. Hutterer (eds.), Vertebrates in the Tropics. Museum Alexander Koenig, Bonn. 424 pp.

1980's

Rickart, E. A., P. D. Heideman, and R. C. B. Utzurrum. 1989. Tent roosting by Scotophilus kuhlii (Chiroptera: Vespertilionidae) in the Philippines. Journal of Tropical Ecology, 5:433-436.
Heideman, P. D., and L. R. Heaney. 1989. Population biology of fruit bats (Pteropodidae) in Philippine submontane rainforest. Journal of Zoology (London), 218:565-586.
Heaney, L. R., P. D. Heideman, E. A. Rickart, R. B. Utzurrum, and J. S. H. Klompen. 1989. Elevational zonation of mammals in the central Philippines. Journal of Tropical Ecology, 5:259-280.
Rickart, E. A., L. R. Heaney, and M. J. Rosenfeld. 1989. Chromosomes of ten species of Philippine fruit bats (Chiroptera: Pteropodidae). Proceedings of the Biological Society of Washington, 102:520-531.
Heideman, P. D. 1989. Temporal and spatial variation in the phenology of flowering and fruiting in a tropical rainforest. Journal of Ecology, 77:1059-1079.
Heideman, P. D. 1989. Delayed development in a paleotropical fruit bat, Haplonycteris fischeri (Pteropodidae). Journal of Reproduction and Fertility, 85:363-382.
Heideman, P. D., and L. R. Heaney. 1989. Population biology of fruit bats (Pteropodidae) in Philippine submontane rainforest. Journal of Zoology (London), 218:564-586.
Heaney. L. R., P. D. Gonzales, and A. C. Alcala. 1988. An annotated checklist of the taxonomic and conservation status of land mammals in the Philippines. Silliman Journal, 34(1987): 32-66.
Heideman, P. D. 1988. The timing of reproduction in the fruit bat Haplonycteris fischeri (Pteropodidae): Geographic variation and delayed development. Journal of Zoology (London), 215:577-595.
Heideman, P. D., L. R. Heaney, R. L. Thomas, and K. R. Erickson. 1987. Patterns of faunal diversity and species abundance of non-volant small mammals on Negros Island, Philippines. Journal of Mammalogy, 68:884-888.
Heaney, L. R., and P. D. Heideman. 1987. Philippine fruit bats, endangered and extinct. Bats, 5:3-5.*
Heideman, P. D., and K. R. Erickson. 1987. The climate and hydrology of the Lake Balinsasayao watershed, Negros Oriental, Philippines. Silliman Journal, 34:82-107.
Heaney, L. R., and A. C. Alcala. 1986. Flat-headed bats (Mammalia, Tylonycteris) from the Philippine Islands. Silliman Journal (Philippines), 33:117-123.
Heaney, L. R. 1986. Biogeography of mammals in Southeast Asia: estimates of rates of colonization, extinction, and speciation. Biological Journal of the Linnean Society, 28:127-165.
Heaney, L. R., and B. D. Patterson. 1986. Introduction. Biological Journal of the Linnean Society, 28:i-iii.
Heaney, L. R., and B. D. Patterson. (eds.) 1986. Island Biogeography of Mammals. Academic Press, London. 271 pp. (initially produced as a special issue of the Biological Journal of the Linnean Society, London).
Heaney, L. R. 1985. Zoogeographic evidence for Middle and Late Pleistocene land bridges to the Philippine Islands. Modern Quaternary Research in Southeast Asia, 9:127-143.
Heaney, L. R. 1985. Systematics of Oriental pygmy squirrels of the genera Exilisciurus and Nannosciurus (Mammalia, Sciuridae). Miscellaneous Publications of the Museum of Zoology, University of Michigan, 170:1-58.
Musser, G. G., L. R. Heaney, and D. S. Rabor. 1985. Philippine rats: description of a new species of Crateromys from Dinagat Island. American Museum Novitates, 2821:1-25.
Musser, G. G., and L. R. Heaney. 1985. Philippine Rattus: a new species from the Sulu Archipelago. American Museum Novitates, 2818:1-32.
Heaney, L. R., and R. L. Peterson. 1984. A new species of tube-nosed fruit bat (Nyctimene) from Negros Island, Philippines (Mammalia: Pteropodidae). Occasional Papers of the Museum of Zoology, University of Michigan, 708:1-16.
Heaney, L. R. 1984. Mammals from Camiguin Island, Philippines. Proceedings of the Biological Society of Washington, 97:119-125.
Heaney, L. R. 1984. Mammalian species richness on islands on the Sunda Shelf, Southeast Asia. Oecologia, 61:11-17.
Heaney, L. R., and R. M. Timm. 1983. Systematics and distribution of shrews of the genus Crocidura (Mammalia: Insectivora) in Vietnam. Proceedings of the Biological Society of Washington, 96:115-120.
Erickson, K. R., and P. D. Heideman. 1983. Notes on the avifauna of the Twin Lakes Region, Negros Oriental, Philippines. Silliman Journal, 30:63-72.
Heaney, L. R., and G. S. Morgan. 1982. A new species of gymnure (Podogymnura) from Dinagat Island, Philippines (Mammalia: Erinaceidae). Proceedings of the Biological Society of Washington, 95:13-26.
Heaney, L. R., and D. S. Rabor. 1982. Mammals of Dinagat and Siargao Island, Philippines. Occasional Papers of the Museum of Zoology, University of Michigan, 699:1-30.
Heaney, L. R., P. D. Heideman, and K. M. Mudar. 1981. Ecological notes on mammals in the Lake Balinsasayao region, Negros Oriental, Philippines. Silliman Journal (Philippines), 28:122-131.
Thorington, R. W., and L. R. Heaney. 1981. Body proportions and gliding adaptations of flying squirrels (Petauristinae). Journal of Mammalogy, 62:101-114.

1970's

Heaney, L. R. 1979. A new species of tree squirrel (Sundasciurus) from Palawan Island, Philippines (Mammalia: Sciuridae). Proceedings of the Biological Society of Washington, 92:280-286.
Heaney, L. R. 1978. Island area and body size of insular mammals: evidence from the tri-colored squirrel (Callosciurus prevostii) of Southeast Asia. Evolution, 32:29-44.